{"id":28605,"date":"2017-04-04T14:16:16","date_gmt":"2017-04-04T14:16:16","guid":{"rendered":"https:\/\/bladmineerders.nl\/animalia\/arthropoda\/insecta\/lepidoptera\/gracillariidae\/dialectica\/dialectica-imperialella\/"},"modified":"2026-07-25T12:47:55","modified_gmt":"2026-07-25T10:47:55","slug":"dialectica-imperialella","status":"publish","type":"page","link":"https:\/\/bladmineerders.nl\/en\/parasites\/animalia\/arthropoda\/insecta\/lepidoptera\/ditrysia\/gracillarioidea\/gracillariidae\/gracillariinae\/dialectica\/dialectica-imperialella\/","title":{"rendered":"Dialectica imperialella"},"content":{"rendered":"<h1><em>Dialectica imperialella<\/em> (Zeller, 1847)<\/h1>\n<p><\/p>\n<h2 class=\"vernacular-name\">fen slender<\/h2>\n<p><\/p>\n<div class=\"beeld\">\n<img loading=\"lazy\" decoding=\"async\" class=\"centreer1\" src=\"https:\/\/bladmineerders.nl\/wp-content\/uploads\/beeld\/_1040_1.jpg\" alt=\"Dialectica imperialella: mine on Symphytum officinale\" width=\"500\" height=\"310\" \/><br \/>\n<br \/>\n<\/p>\n<p class=\"en-bijschrift\"><em>Symphytum officinale<\/em>, Belgium, prov. Li\u00e8ge, Oreye: mine with young, colourless, larva \u00a9 Jean-Yves Baugn\u00e9e<\/p>\n<p>\n<\/p><\/div>\n<p><!-- beeld --><\/p>\n<div class=\"beeld\">\n<img loading=\"lazy\" decoding=\"async\" class=\"centreer1\" src=\"https:\/\/bladmineerders.nl\/wp-content\/uploads\/beeld\/9639_up.jpg\" alt=\"Dialectica imperialella: mine on Symphytum officinale\" width=\"500\" height=\"143\" \/><br \/>\n<br \/>\n<\/p>\n<p class=\"en-bijschrift\"><em>Symphytum officinale<\/em>, Nieuwendam; mine, upper side<\/p>\n<p>\n<\/p><\/div>\n<p><!-- beeld --><\/p>\n<div class=\"beeld\">\n<img loading=\"lazy\" decoding=\"async\" class=\"centreer1\" src=\"https:\/\/bladmineerders.nl\/wp-content\/uploads\/beeld\/9639_low.jpg\" alt=\"Dialectica imperialella: mine on Symphytum officinale\" width=\"500\" height=\"136\" \/><br \/>\n<br \/>\n<\/p>\n<p class=\"en-bijschrift\">same mine, lower side<\/p>\n<p>\n<\/p><\/div>\n<p><!-- beeld --><\/p>\n<div class=\"beeld\">\n<img loading=\"lazy\" decoding=\"async\" class=\"centreer1\" src=\"https:\/\/bladmineerders.nl\/wp-content\/uploads\/beeld\/9639_tr.jpg\" alt=\"Dialectica imperialella: mine on Symphytum officinale\" width=\"500\" height=\"213\" \/><br \/>\n<br \/>\n<\/p>\n<p class=\"en-bijschrift\">same mine again, lighted from behind, with full grown larva<\/p>\n<p>\n<\/p><\/div>\n<p><!-- beeld --><\/p>\n<div class=\"beeld\">\n<img loading=\"lazy\" decoding=\"async\" class=\"centreer1\" src=\"https:\/\/bladmineerders.nl\/wp-content\/uploads\/beeld\/22955_3.jpg\" alt=\"Dialectica imperialella: detail of  start of mine on Symphytum officinale\" width=\"500\" height=\"321\" \/><br \/>\n<br \/>\n<\/p>\n<p class=\"en-bijschrift\">Symphytum officinale, Nieuwendam: detail detail of the start of the mine.<\/p>\n<p>\n<\/p><\/div>\n<p><!-- beeld --><\/p>\n<div class=\"beeld\">\n<img loading=\"lazy\" decoding=\"async\" class=\"centreer1\" src=\"https:\/\/bladmineerders.nl\/wp-content\/uploads\/beeld\/_1290_7.jpg\" alt=\"Dialectica imperialella: mine on Pentaglottis sempervirens\" width=\"500\" height=\"375\" \/><br \/>\n<br \/>\n<\/p>\n<p class=\"en-bijschrift\"><em>Pentaglottis sempervirens<\/em>, Belgium, West Flanders, Wielsbeke \u00a9 Steve Wullaert<\/p>\n<p>\n<\/p><\/div>\n<p><!-- beeld --><\/p>\n<div class=\"beeld\">\n<img loading=\"lazy\" decoding=\"async\" class=\"centreer1\" src=\"https:\/\/bladmineerders.nl\/wp-content\/uploads\/beeld\/_1290_9.jpg\" alt=\"Dialectica imperialella: mine on Pentaglottis sempervirens\" width=\"500\" height=\"375\" \/><br \/>\n<br \/>\n<\/p>\n<p class=\"en-bijschrift\">underside of the leaf; exceptionally heavy infestation<\/p>\n<p>\n<\/p><\/div>\n<p><!-- beeld --><\/p>\n<div class=\"beeld\">\n<img loading=\"lazy\" decoding=\"async\" class=\"centreer1\" src=\"https:\/\/bladmineerders.nl\/wp-content\/uploads\/beeld\/_1587_11.jpg\" alt=\"Dialectica imperialella: mine on Echium plantagineum\" width=\"500\" height=\"169\" \/><br \/>\n<br \/>\n<\/p>\n<p class=\"en-bijschrift\"><em>Echium plantagineum<\/em>, Belgium, prov Antwerp, Mol; \u00a9 Carina Van Steenwinkel<\/p>\n<p>\n<\/p><\/div>\n<p><!-- beeld --><\/p>\n<p><br \/>\n<\/p>\n<h3 class=\"head-mine\">mine<\/h3>\n<p>A number of eggs are deposited over a wide area of the underside of a leaf. The larvae make very wide epidermal galleries, that are hardly discernable, but for the tread thin line of frass that is glued to the floor of the mine. Gradually the larvae come together, and the mine becomes deeper, and finally becoming locally full depth. Only then the mine is apparent from above as some disconnected brownish spots. At the underside the mine is quite larger, membraneous. Pupation usually is outside the mine.<\/p>\n<p><\/p>\n<p><br \/>\n<\/p>\n<h3 class=\"head-hostplant\">hostplants<\/h3>\n<p>Boraginaceae, oligophagous<\/p>\n<p><\/p>\n<p class=\"latinnames\"><a href=\"https:\/\/bladmineerders.nl\/buglossoides\">Aegonychon<\/a> purpurocaeruleum; <a href=\"https:\/\/bladmineerders.nl\/lithospermum\">Lithospermum<\/a> officinale; <a href=\"https:\/\/bladmineerders.nl\/pulmonaria\">Pulmonaria<\/a> angustifolia, officinalis; <a href=\"https:\/\/bladmineerders.nl\/symphytum\">Symphytum<\/a> officinale.<\/p>\n<p><br \/>\n<\/p>\n<p>Found in 2012 by Steve Wullaert on <em><a href=\"https:\/\/bladmineerders.nl\/pentaglottis\">Pentaglottis<\/a> sempervirens<\/em>, by Carina Van Steenwinkel on <em><a href=\"https:\/\/bladmineerders.nl\/echium\">Echium<\/a> plantagineum<\/em>.<\/p>\n<p><\/p>\n<p><br \/>\n<\/p>\n<h3 class=\"head-phenology\">phenology<\/h3>\n<p>Larvae in August &#8211; September (Emmet, Watkinson &#038; Wilson, 1985a).<\/p>\n<p><\/p>\n<p><br \/>\n<\/p>\n<h3 class=\"head-distribution\">BENELUX<\/h3>\n<p>BE recorded (Phegea, 2009).<\/p>\n<p>NE recorded (Kuchlein &#038; de Vos, 1999a; Microlepidoptera.nl, 2009).<\/p>\n<p>LUX recorded (Fauna Europaea, 2009).<\/p>\n<p><\/p>\n<p><br \/>\n<\/p>\n<h3 class=\"head-distribution\">distribution within Europe<\/h3>\n<p>From Denmark to the Pyrenees, Italy, and Romania, and from the UK to Russia and Ukraine (Fauna Europaea, 2009).<\/p>\n<p><\/p>\n<p><br \/>\n<\/p>\n<h3 class=\"topic\"><a href=\"https:\/\/bladmineerders.nl\/dialectica-imperialella-larva\/\">larva<\/a><\/h3>\n<p><\/p>\n<p><br \/>\n<\/p>\n<h3 class=\"head-pupa\">pupa<\/h3>\n<p>Described by Pato\u010dka (2001b), Pato\u010dka &#038; Tur\u010d\u00e1ni (2005a).<\/p>\n<p><\/p>\n<p><br \/>\n<\/p>\n<h3 class=\"head-synonym\">synonyms<\/h3>\n<p><em>Acrocercops, Gracillaria imperialella<\/em>.<\/p>\n<p><\/p>\n<h3 class=\"head-references\">references<\/h3>\n<p class=\"references\">Barton (2018a), Beiger (1980a), Bengtsson &#038; Johansson (2011a), Biesenbaum (2010a), Buszko (1987a, 1992b), Doets (1946a), Dr\u0103ghia (1968a), Emmet, Watkinson &#038; Wilson (1985a), Falck (2025b), Gomboc &#038; Kirichenko (2022a), Huisman, Koster, van Nieukerken &#038; Ulenberg (2001a, 2003a, 2004a, 2005a, 2006a), Kasy (1965a), Kuchlein &#038; Donner (1993a), Klimesch (1950c), Kuchlein &#038; Donner (1993a), Kuchlein &#038; de Vos (1999a), Mifsud &#038; Askew (2019a), van Nieukerken, Gielis, Huisman ao (1993a), Pato\u010dka (2001b), Pato\u010dka &#038; Tur\u010d\u00e1ni (2005a), De Prins (2010a), De Prins &#038; Steeman (2013a), De Prins, Steeman &#038; Sierens (2016a), Star\u00fd (1930a), Sz\u0151cs (1977a), Wieser &#038; Huemer (1999a).<\/p>","protected":false},"excerpt":{"rendered":"<p>Dialectica imperialella (Zeller, 1847) fen slender Symphytum officinale, Belgium, prov. Li\u00e8ge, Oreye: mine with young, colourless, larva \u00a9 Jean-Yves Baugn\u00e9e Symphytum officinale, Nieuwendam; mine, upper side same mine, lower side same mine again, lighted from behind, with full grown larva Symphytum officinale, Nieuwendam: detail detail of the start of the mine. Pentaglottis sempervirens, Belgium, West [&hellip;]<\/p>\n","protected":false},"author":1,"featured_media":0,"parent":28603,"menu_order":0,"comment_status":"closed","ping_status":"closed","template":"parasite-species.php","meta":{"footnotes":""},"categories":[7],"tags":[],"class_list":["post-28605","page","type-page","status-publish","hentry","category-parasite"],"_links":{"self":[{"href":"https:\/\/bladmineerders.nl\/en\/wp-json\/wp\/v2\/pages\/28605","targetHints":{"allow":["GET"]}}],"collection":[{"href":"https:\/\/bladmineerders.nl\/en\/wp-json\/wp\/v2\/pages"}],"about":[{"href":"https:\/\/bladmineerders.nl\/en\/wp-json\/wp\/v2\/types\/page"}],"author":[{"embeddable":true,"href":"https:\/\/bladmineerders.nl\/en\/wp-json\/wp\/v2\/users\/1"}],"replies":[{"embeddable":true,"href":"https:\/\/bladmineerders.nl\/en\/wp-json\/wp\/v2\/comments?post=28605"}],"version-history":[{"count":3,"href":"https:\/\/bladmineerders.nl\/en\/wp-json\/wp\/v2\/pages\/28605\/revisions"}],"predecessor-version":[{"id":990633,"href":"https:\/\/bladmineerders.nl\/en\/wp-json\/wp\/v2\/pages\/28605\/revisions\/990633"}],"up":[{"embeddable":true,"href":"https:\/\/bladmineerders.nl\/en\/wp-json\/wp\/v2\/pages\/28603"}],"wp:attachment":[{"href":"https:\/\/bladmineerders.nl\/en\/wp-json\/wp\/v2\/media?parent=28605"}],"wp:term":[{"taxonomy":"category","embeddable":true,"href":"https:\/\/bladmineerders.nl\/en\/wp-json\/wp\/v2\/categories?post=28605"},{"taxonomy":"post_tag","embeddable":true,"href":"https:\/\/bladmineerders.nl\/en\/wp-json\/wp\/v2\/tags?post=28605"}],"curies":[{"name":"wp","href":"https:\/\/api.w.org\/{rel}","templated":true}]}}