{"id":32244,"date":"2017-02-15T13:02:54","date_gmt":"2017-02-15T13:02:54","guid":{"rendered":"https:\/\/bladmineerders.nl\/host-plants\/spermatopsida\/angiosperma\/fabids\/rosales\/maloideae\/crataegus\/"},"modified":"2025-10-18T10:13:40","modified_gmt":"2025-10-18T08:13:40","slug":"crataegus","status":"publish","type":"page","link":"https:\/\/bladmineerders.nl\/en\/host-plants\/plantae\/spermatopsida\/angiosperma\/eudicots\/superrosids\/rosids\/fabids\/rosales\/rosaceae\/amygdaloideae\/maleae\/crataegus\/","title":{"rendered":"Crataegus"},"content":{"rendered":"<p><\/p>\n<h1><em>Crataegus<\/em> hawthorn<\/h1>\n<p class=\"remark\"> Incl. <em>x Crataemespilus<\/em>.<\/p>\n<h4>(For a dichotomous table for galls on <em>Crataegus<\/em> by Hans Roskam click <a href=\"https:\/\/bladmineerders.nl\/gallers-on-crataegus\/\">here<\/a>)<\/p>\n<h4>\n<h2>Dichotomous table for leafminers<\/h2>\n<div class=\"couplet\">\n<p><strong>1a<\/strong> mine in fallen leaves: <a href=\"https:\/\/bladmineerders.nl\/neocoenorrhinus-pauxillus\"><em>Neocoenorrhinus pauxillus<\/em><\/a><\/p>\n<p><strong>1b<\/strong> round excisions, 3-4 mm diam.: <a href=\"https:\/\/bladmineerders.nl\/incurvaria-masculella\"><em>Incurvaria masculella<\/em><\/a><\/p>\n<p><strong>1c<\/strong> tentiform mine, &gt; 1 cm =&gt; 2<\/p>\n<p><strong>1d<\/strong> small, &lt; 1 cm, blotch or corridor =&gt; 6<\/p>\n<p><strong>1e<\/strong> blotch, &gt; 1 cm, without recognisable preceding corridor =&gt; 14<\/p>\n<p><strong>1f<\/strong> corridor, &gt; 1 cm, not strongly widened as the end =&gt; 20<\/p>\n<p><strong>1g<\/strong> corridor, &gt; 1 cm, strongly widened at the end, or forming a secondary blotch =&gt; 22<\/p>\n<p><strong>1h<\/strong> fleck mine =&gt; 29<\/p>\n<p><strong>1i<\/strong> galls, etc =&gt; <a href=\"https:\/\/bladmineerders.nl\/crataegus\/#all\">Tables for all parasites per species<\/a><\/p>\n<\/div>\n<p><!-- couplet --><\/p>\n<div class=\"couplet\">\n<p><strong>2a<\/strong> mine upper-surface, silvery =&gt; 3<\/p>\n<p><strong>2b<\/strong> mine lower-surface =&gt; 4<\/p>\n<\/div>\n<p><!-- couplet --><\/p>\n<div class=\"couplet\">\n<p><strong>3a<\/strong> fully developed mine silvery white, without black speckles: <a href=\"https:\/\/bladmineerders.nl\/phyllonorycter-leucographella\"><em>Phyllonorycter leucographella<\/em><\/a><\/p>\n<p><strong>3b<\/strong> fully developed mine dirty white because of numerous black speckles: <a href=\"https:\/\/bladmineerders.nl\/phyllonorycter-corylifoliella\"><em>Phyllonorycter corylifoliella<\/em><\/a><\/p>\n<\/div>\n<p><!-- couplet --><\/p>\n<div class=\"couplet\">\n<p><strong>4a<\/strong> lower epidermis brownish, wrinkled but not strongly folded; mine roundish, in a leaf lobe: <a href=\"https:\/\/bladmineerders.nl\/parornix-anglicella\"><em>Parornix anglicella<\/em><\/a><\/p>\n<p><strong>4b<\/strong> lower epidermis yellow-green, strongly folded; mine elongated along the leaf margin or between two veins =&gt; 5<\/p>\n<\/div>\n<p><!-- couplet --><\/p>\n<div class=\"couplet\">\n<p><strong>5a<\/strong> pupa grey-brown, without, or in a very thin, yellow cocoon (very common on Hawthorn): <a href=\"https:\/\/bladmineerders.nl\/phyllonorycter-oxyacanthae\"><em>Phyllonorycter oxyacanthae<\/em><\/a><\/p>\n<p><strong>5b<\/strong> pupa reddish-brown or yellowish, in a white cocoon:<em><a href=\"https:\/\/bladmineerders.nl\/phyllonorycter-mespilella\">Phyllonorycter mespilella<\/a><\/em><\/p>\n<\/div>\n<p><!-- couplet --><\/p>\n<div class=\"couplet\">\n<p><strong>6a<\/strong> drop- or pear-shaped blotch, for its greatest part filled with frass =&gt; 7<\/p>\n<p><strong>6b<\/strong> not this shape, or not that much frass =&gt; 8<\/p>\n<\/div>\n<p><!-- couplet --><\/p>\n<div class=\"couplet\">\n<p><strong>7a<\/strong> larvae in May-June: <a href=\"https:\/\/bladmineerders.nl\/rhamphus-subaeneus\"><em>Rhamphus subaeneus<\/em><\/a><\/p>\n<p><strong>7b<\/strong> larvae in September-October: <a href=\"https:\/\/bladmineerders.nl\/rhamphus-oxyacanthae\"><em>Rhamphus oxyacanthae<\/em><\/a><\/p>\n<\/div>\n<p><!-- couplet --><\/p>\n<div class=\"couplet\">\n<p><strong>8a<\/strong> very clear corridor or blotch of max. 3 mm, often several in a leaf =&gt; 9<\/p>\n<p><strong>8b<\/strong> mine less transparent and\/or larger; no more than 1 in a leaf =&gt; 11<\/p>\n<\/div>\n<p><!-- couplet --><\/p>\n<div class=\"couplet\">\n<p><strong>9a<\/strong> larva cuts upper and lower epidermis out of the mine, and drops to the ground: <a href=\"https:\/\/bladmineerders.nl\/incurvaria-masculella\"><em>Incurvaria masculella<\/em><\/a><\/p>\n<p><strong>9b<\/strong> larva doesn&#8217;t make an excision =&gt; 10<\/p>\n<\/div>\n<p><!-- couplet --><\/p>\n<div class=\"couplet\">\n<p><strong>10a<\/strong> larva: head golden brown: <a href=\"https:\/\/bladmineerders.nl\/paraswammerdamia-nebulella\"><em>Paraswammerdamia nebulella<\/em><\/a><\/p>\n<p><strong>10b<\/strong> head black with brown and white lines: <a href=\"https:\/\/bladmineerders.nl\/scythropia-crataegella\"><em>Scythropia crataegella<\/em><\/a><\/p>\n<\/div>\n<p><!-- couplet --><\/p>\n<div class=\"couplet\">\n<p><strong>11a<\/strong> sides of the mine eaten out irregularly; mine often branched: <a href=\"https:\/\/bladmineerders.nl\/recurvaria-nanella\"><em>Recurvaria nanella<\/em><\/a><\/p>\n<p><strong>11b<\/strong>sides smooth; mine usually unbranched =&gt; 12<\/p>\n<\/div>\n<p><!-- couplet --><\/p>\n<div class=\"couplet\">\n<p><strong>12a<\/strong> mine with much frass; a hooklike corridor, mostly in a vein axil: <a href=\"https:\/\/bladmineerders.nl\/bucculatrix-bechsteinella\"><em>Bucculatrix bechsteinella<\/em><\/a><\/p>\n<p><strong>12b<\/strong> mine with little or no frass; irregularly shaped  =&gt; 13<\/p>\n<\/div>\n<p><!-- couplet --><\/p>\n<div class=\"couplet\">\n<p><strong>13a<\/strong> larva: pinacula colourless (the bases of the setae themselves are black) : <em><a href=\"https:\/\/bladmineerders.nl\/cnephasia-incertana\">Cnephasia incertana<\/a><\/em><\/p>\n<p><strong>13b<\/strong> pinacula black: <em><a href=\"https:\/\/bladmineerders.nl\/cnephasia-asseclana\">Cnephasia asseclana<\/a><\/em><\/p>\n<\/div>\n<p><!-- couplet --><\/p>\n<div class=\"couplet\">\n<p><strong>14a<\/strong> mine lower-surface, grey, later brown: youth mine of <a href=\"https:\/\/bladmineerders.nl\/parornix-anglicella\"><em>Parornix anglicella<\/em><\/a><\/p>\n<p><strong>14b<\/strong> mine upper-surface, silvery, elongated, positioned over a thick vein: youth mine of <a href=\"https:\/\/bladmineerders.nl\/phyllonorycter-corylifoliella\"><em>Phyllonorycter corylifoliella<\/em><\/a> or <a href=\"https:\/\/bladmineerders.nl\/phyllonorycter-leucographella\"><em>Ph. leucographella<\/em><\/a><\/p>\n<p><strong>14c<\/strong> mine full depth =&gt; 15<\/p>\n<\/div>\n<p><!-- couplet --><\/p>\n<div class=\"couplet\">\n<p><strong>15a<\/strong> oviposition site (usually at the tip of a leaf lobe) covered by a shining black drop of dried secretion: <a href=\"https:\/\/bladmineerders.nl\/trachys-minutus\"><em>Trachys minutus<\/em><\/a><\/p>\n<p><strong>15b<\/strong> no such drop =&gt; 16<\/p>\n<\/div>\n<p><!-- couplet --><\/p>\n<div class=\"couplet\">\n<p><strong>16a<\/strong> mine triangular, between midrib and a lateral vein; frass warm-brown: <a href=\"https:\/\/bladmineerders.nl\/stigmella-perpygmaeella\"><em>Stigmella perpygmaeella<\/em><\/a><\/p>\n<p><strong>16b<\/strong> mine round or irregular, frass grey or black =&gt; 17<\/p>\n<\/div>\n<p><!-- couplet --><\/p>\n<div class=\"couplet\">\n<p><strong>17a<\/strong> larva (always present) pear-shaped: <a href=\"https:\/\/bladmineerders.nl\/rhamphus-oxyacanthae\"><em>Rhamphus oxyacanthae<\/em><\/a><\/p>\n<p><strong>17b<\/strong> larva (if present) slender =&gt; 18<\/p>\n<\/div>\n<p><!-- couplet --><\/p>\n<div class=\"couplet\">\n<p><strong>18a<\/strong> mine an irregular blotch; elsewhere on the leaf, or on neighbouring leaves, long slender galleries with a central frass line: <em><a href=\"https:\/\/bladmineerders.nl\/lyonetia-prunifoliella\">Lyonetia prunifoliella<\/a><\/em><\/p>\n<p><strong>18b<\/strong> circular blotch; not associated with gallery mines =&gt; 19<\/p>\n<\/div>\n<p><!-- couplet --><\/p>\n<div class=\"couplet\">\n<p><strong>19a<\/strong> mine in the centre of the leaf; frass in concentric circles: <a href=\"https:\/\/bladmineerders.nl\/leucoptera-malifoliella\"><em>Leucoptera malifoliella<\/em><\/a><\/p>\n<p><strong>19b<\/strong> mine in a leaf lobe; frass concentrated in the centre, not in circles: <a href=\"https:\/\/bladmineerders.nl\/stigmella-paradoxa\"><em>Stigmella paradoxa<\/em><\/a><\/p>\n<\/div>\n<p><!-- couplet --><\/p>\n<div class=\"couplet\">\n<p><strong>20a<\/strong> corridor begins at a shining globular egg shell; frass coiled: <a href=\"https:\/\/bladmineerders.nl\/stigmella-oxyacanthella\"><em>Stigmella oxyacanthella<\/em><\/a><\/p>\n<p><strong>20b<\/strong> no egg shell; frass in a narrow central line =&gt; 21<\/p>\n<\/div>\n<p><!-- couplet --><\/p>\n<div class=\"couplet\">\n<p><strong>21a<\/strong> elsehwere on the leaf, or on leaves nearby, large, full depth blotch mines (sometimes connected to the corridor): <em><a href=\"https:\/\/bladmineerders.nl\/lyonetia-prunifoliella\">Lyonetia prunifoliella<\/a><\/em><\/p>\n<p><strong>21b<\/strong> no associated blotch mines: <a href=\"https:\/\/bladmineerders.nl\/lyonetia-clerkella\"><em>Lyonetia clerkella<\/em><\/a><\/p>\n<\/div>\n<p><!-- couplet --><\/p>\n<div class=\"couplet\">\n<p><strong>22a<\/strong> frass in the widened part of the mine, at least partly, coiled; frass warm-brown =&gt; 23<\/p>\n<p><strong>22b<\/strong> frass in the widened part of the mine not coiled, black or brown-black (frass may be reddish-brown in the initial corridor, and be coiled there) =&gt; 24<\/p>\n<\/div>\n<p><!-- couplet --><\/p>\n<div class=\"couplet\">\n<p><strong>23a<\/strong> mine triangular, between two lateral veins, or between a lateral vein and the leaf margin; initial corridor largely overrun by the secondary blotch; larva yellow: <a href=\"https:\/\/bladmineerders.nl\/stigmella-perpygmaeella\"><em>Stigmella perpygmaeella<\/em><\/a><\/p>\n<p><strong>23b<\/strong> mine tresspassing over the lateral veins, initial corridor not overrun; larva green: <a href=\"https:\/\/bladmineerders.nl\/stigmella-crataegella\"><em>Stigmella crataegella<\/em><\/a><\/p>\n<\/div>\n<p><!-- couplet --><\/p>\n<div class=\"couplet\">\n<p><strong>24a<\/strong> mine begins upon the petiole =&gt; 25<\/p>\n<p><strong>24b<\/strong> mine begins in the leaf blade =&gt; 26<\/p>\n<\/div>\n<p><!-- couplet --><\/p>\n<div class=\"couplet\">\n<p><strong>25a<\/strong> frass in the entire mine brown-black or black; larva whitish: <a href=\"https:\/\/bladmineerders.nl\/ectoedemia-atricollis\"><em>Ectoedemia atricollis<\/em><\/a><\/p>\n<p><strong>25b<\/strong> frass red-brown in the initial corridor, black in the secondary blotch; larva yellow: <a href=\"https:\/\/bladmineerders.nl\/stigmella-regiella\"><em>Stigmella regiella<\/em><\/a><\/p>\n<\/div>\n<p><!-- couplet --><\/p>\n<div class=\"couplet\">\n<p><strong>26a<\/strong> initial corrior wholly or largely free from the leaf margin, with black frass; larva with pale brown head =&gt; 27<\/p>\n<p><strong>26b<\/strong> initial corridor follows the leaf margin for most of its length; frass there brown or black =&gt; 28<\/p>\n<\/div>\n<p><!-- couplet --><\/p>\n<div class=\"couplet\">\n<p><strong>27a<\/strong> Eastern Mediterranean species on <em>Crataegus azarolus<\/em>: <em><a href=\"https:\/\/bladmineerders.nl\/stigmella-azaroli\">Stigmella azaroli<\/a><\/em><\/p>\n<p><strong>27b<\/strong> pan-European species on different Hawthorn species: <a href=\"https:\/\/bladmineerders.nl\/stigmella-hybnerella\"><em>Stigmella hybnerella<\/em><\/a><\/p>\n<\/div>\n<p><!-- couplet --><\/p>\n<div class=\"couplet\">\n<p><strong>28a<\/strong> in the initial corridor the frass is black; if the first part of the corridor is free from the leaf margin it twists around the oviposition spot; larva whitish with black-brown head: <a href=\"https:\/\/bladmineerders.nl\/ectoedemia-atricollis\"><em>Ectoedemia atricollis<\/em><\/a><\/p>\n<p><strong>28b<\/strong> in the initial corridor the frass is brown; initial corridor not twisted; larva yellow with pale brown head: <a href=\"https:\/\/bladmineerders.nl\/stigmella-regiella\"><em>Stigmella regiella<\/em><\/a><\/p>\n<\/div>\n<p><!-- couplet --><\/p>\n<div class=\"couplet\">\n<p><strong>29a<\/strong> pistol case: <a href=\"https:\/\/bladmineerders.nl\/coleophora-anatipenella\"><em>Coleophora anatipenella<\/em><\/a><\/p>\n<p><strong>29b<\/strong> lobe case: <a href=\"https:\/\/bladmineerders.nl\/coleophora-violacea\"><em>Coleophora violacea<\/em><\/a><\/p>\n<p><strong>29c<\/strong> tubular silken case: <a href=\"https:\/\/bladmineerders.nl\/coleophora-trigeminella\"><em>Coleophora trigeminella<\/em><\/a><\/p>\n<p><strong>29d<\/strong> composite leaf case =&gt; 30<\/p>\n<p><strong>29e<\/strong> tubular leaf case =&gt; 31<\/p>\n<\/div>\n<p><!-- couplet --><\/p>\n<div class=\"couplet\">\n<p><strong>30a<\/strong> rear end of the case crooked, like the handle of a walking stick: <a href=\"https:\/\/bladmineerders.nl\/coleophora-hemerobiella\"><em>Coleophora hemerobiella<\/em><\/a><\/p>\n<p><strong>30b<\/strong> rear end at most at right angle with the rest of the case: <a href=\"https:\/\/bladmineerders.nl\/coleophora-spinella\"><em>Coleophora spinella<\/em><\/a><\/p>\n<\/div>\n<p><!-- couplet --><\/p>\n<div class=\"couplet\">\n<p><strong>31a<\/strong> case with large irregular extensions: <a href=\"https:\/\/bladmineerders.nl\/coleophora-siccifolia\"><em>Coleophora siccifolia<\/em><\/a><\/p>\n<p><strong>31b<\/strong> case without such extensions =&gt; 32<\/p>\n<\/div>\n<p><!-- couplet --><\/p>\n<div class=\"couplet\">\n<p><strong>32a<\/strong> case 8-11 mm; mouth angle 90\u00b0: <a href=\"https:\/\/bladmineerders.nl\/coleophora-hemerobiella\"><em>Coleophora hemerobiella<\/em><\/a><\/p>\n<p><strong>32b<\/strong> case 6-7 mm; mouth angle 60\u00b0 =&gt; 33<\/p>\n<\/div>\n<p><!-- couplet --><\/p>\n<div class=\"couplet\">\n<p><strong>33a<\/strong> SW Mediterranean species: <em><a href=\"https:\/\/bladmineerders.nl\/coleophora-chiclanensis\">Coleophora chiclanensis<\/a><\/em><\/p>\n<p><strong>33b<\/strong> pan-European species: <a href=\"https:\/\/bladmineerders.nl\/coleophora-spinella\"><em>Coleophora spinella<\/em><\/a> (most common on Hawthorn), but see also <em><a href=\"https:\/\/bladmineerders.nl\/coleophora-prunifoliae\">C. prunifoliae<\/a><\/em> and <em><a href=\"https:\/\/bladmineerders.nl\/coleophora-serratella\">C serratella<\/a><\/em>.<\/p>\n<\/div>\n<p><!-- couplet --><\/p>\n<p>Not included in the key: <em><a href=\"https:\/\/bladmineerders.nl\/coleophora-adjectella\">Coleophora adjectella<\/a><\/em> (contradictions in the description), <em><a href=\"https:\/\/bladmineerders.nl\/coleophora-coracipennella\">Coleophora coracipennella<\/a><\/em>, <em><a href=\"https:\/\/bladmineerders.nl\/coleophora-potentillae\">Coleophora potentillae<\/a><\/em>l <em><a href=\"https:\/\/bladmineerders.nl\/liriomyza-trifolii\">Liriomyza trifolii<\/a><\/em>; <a href=\"https:\/\/bladmineerders.nl\/phyllonorycter-blancardella\"><em>Phyllonorycter blancardella<\/em><\/a> (xenophagy?), <em><a href=\"https:\/\/bladmineerders.nl\/phyllonorycter-macedonica\">Phyllonorycter macedonica<\/a><\/em> (unsufficiently described); <em><a href=\"https:\/\/bladmineerders.nl\/phyllonorycter-sorbi\">Phyllonorycter sorbi<\/a><\/em>; <em><a href=\"https:\/\/bladmineerders.nl\/stigmella-mespilicola\">Stigmella mespilicola<\/a><\/em> (occurence on this host unconfirmed).<\/p>\n<p><\/p>","protected":false},"excerpt":{"rendered":"<p>Crataegus hawthorn Incl. x Crataemespilus. (For a dichotomous table for galls on Crataegus by Hans Roskam click here) Dichotomous table for leafminers 1a mine in fallen leaves: Neocoenorrhinus pauxillus 1b round excisions, 3-4 mm diam.: Incurvaria masculella 1c tentiform mine, &gt; 1 cm =&gt; 2 1d small, &lt; 1 cm, blotch or corridor =&gt; 6 [&hellip;]<\/p>\n","protected":false},"author":1,"featured_media":0,"parent":294096,"menu_order":0,"comment_status":"closed","ping_status":"closed","template":"host-genus.php","meta":{"footnotes":""},"categories":[6],"tags":[],"class_list":["post-32244","page","type-page","status-publish","hentry","category-host-plant"],"_links":{"self":[{"href":"https:\/\/bladmineerders.nl\/en\/wp-json\/wp\/v2\/pages\/32244","targetHints":{"allow":["GET"]}}],"collection":[{"href":"https:\/\/bladmineerders.nl\/en\/wp-json\/wp\/v2\/pages"}],"about":[{"href":"https:\/\/bladmineerders.nl\/en\/wp-json\/wp\/v2\/types\/page"}],"author":[{"embeddable":true,"href":"https:\/\/bladmineerders.nl\/en\/wp-json\/wp\/v2\/users\/1"}],"replies":[{"embeddable":true,"href":"https:\/\/bladmineerders.nl\/en\/wp-json\/wp\/v2\/comments?post=32244"}],"version-history":[{"count":0,"href":"https:\/\/bladmineerders.nl\/en\/wp-json\/wp\/v2\/pages\/32244\/revisions"}],"up":[{"embeddable":true,"href":"https:\/\/bladmineerders.nl\/en\/wp-json\/wp\/v2\/pages\/294096"}],"wp:attachment":[{"href":"https:\/\/bladmineerders.nl\/en\/wp-json\/wp\/v2\/media?parent=32244"}],"wp:term":[{"taxonomy":"category","embeddable":true,"href":"https:\/\/bladmineerders.nl\/en\/wp-json\/wp\/v2\/categories?post=32244"},{"taxonomy":"post_tag","embeddable":true,"href":"https:\/\/bladmineerders.nl\/en\/wp-json\/wp\/v2\/tags?post=32244"}],"curies":[{"name":"wp","href":"https:\/\/api.w.org\/{rel}","templated":true}]}}