Clematis traveller’s joy
Dichotomous table for leafminers
2a larva a maggot; on Clematis cirrhosa: Phytomyza cirrhosae
2b larva with thoracic feet and chitinised head; on other hostplants => 3
3a larva with prolegs; mine full depth, very transparent => 4
3b no prolegs; mine not entirely full depth, greenish when lighted from behind => 5
4a on Clematis alpina: Pseudodineura clematidis
4b on Clematis recta: Ps. clematidisrectae
5a eastern European species: Argopus nigritarsis
5b more a species of central and western Europe: Argopus ahrensii, brevis
6a corridor short, ending upon a thick vein or into the leaf base => 7
6b corridor much longer => 8
7a on Clematis flammula or recta: Ptochomyza mayeri
7b on othe species: Phytomyza clematidis
8a mine begins wth a densely wound, intestine-like, upper-surface corridor: Phytomyza anemones
8b mine begins with very long and narrow lower-surface part, that several times crosses the leaf: Phytomyza kaltenbachi
8c mine begins with a knot or spiral, or a short lower-surface corridor (may be along the midrib) => 9
9a lower-surface section mostly a densely wound spiral => 10
9b lower-surface section a corridor of a few mm => 12
10a frass in two rows; corridor not associated with the leaf margin; northern Europe and the mountains of central Europe: Phytomyza pulsatillae
10b frass in pearl chains of strings at one side of the corridor; parts of the corridor follow the leaf margin; south and central European lowlands => 11
11a lower-surface section a more or less tortuous, epidermal, silvery corridor with a fine black frass line: Phytomyza dalmatiensis
11b lower-surface section a dense knot in the sponge parenchyma, finally forming a small full depth transparent patch: Phytomyza rectae
12a frass in long threads that over long tracks lie at the same side of the corridor; mine usually begins in the tip of a leaflet; in upper-surface part of the mine the corridor sides are smooth: Phytomyza vitalbae
12b frass in pearl chains, alternating along the sides of the mine, sometimes almost seeming central; mine less often beginning in the tip of a leaflet; corridor sides in upper-surface part of the mine irregularly eaten out: Phytomyza fulgens
Not included in the key: Phytomyza actaeae.