Echium viper’s bugloss
(For a dichotomous table for galls on Echium by Hans Roskam click here)
Dichotomous table for leafminers
1a fleck mine => 2
1b mine full depth => 3
1c mine entirely or largely upper- or lower-surface => 7
1d galls, etc => Tables for all parasites per species
Dichotomous table for leafminers
1a fleck mine => 2
1b mine full depth => 3
1c mine entirely or largely upper- or lower-surface => 7
1d galls, etc => Tables for all parasites per species
2a larva in a helicoidal case: Apterona helicoidella
2b larva in a tubular case: Coleophora pennella
3a mine large with a strongly inflated centre => 4
3b mine small, irregular, without a clear centre => 5
4a mine mainly in the lower part of the leaf, causing strong malformation; often on Echium: Cynaeda dentalis
4b mine usually in the top half of the leaf, not causing strong malformation; rarely on Echium: Epascestria pustulalis
5a larva: pinacula colourless (the base of the setae is black): Cnephasia incertana
5b pinacula black => 6
6a larva: last abdomen segment with a black anal comb: Cnephasia asseclana
6b no anal comb present: Cnephasia stephensiana
7a larva with feet and a sclerotised head => 8
7b larva a maggot => 10
8a pupa naked in the mine, works itself halfway out of the mine prior to emergence: Dialectica scalariella
8b pupa remains in the mine => 9
9a pupa with a dense cover of fine, short “hairs”: Aproaerema lerauti
9b pop with a smooth integument: Perittia echiella
10a gallery from begin to end; pupation within the mine; frass in isolated grains: Chromatomyia horticola
10b blotch, may be preceded by a corridor; frass in thread fragments => 11
11a mine begins with a long an slender corridor part (often overrun by the later blotch, but as a rule still recognisable in the frass pattern); cephalic skeleton of the larva with three rearwards directed arms:: Agromyza abiens
11b no initial corridor; cephalic skeleton with two rearwards pointing arms: Phytomyza medicaginis
Not included in the key: Coleophora tauricella (inadequately described); Dialectica imperialella.