Symphytum comfrey
(For a dichotomous table for galls on Symphytum by Hans Roskam click here)
Dichotomous table for leafminers
1a fleck mine; larva mines from a case => 2
1b corridor or blotch mine; larva mines from the inside => 3
1c galls, etc. => Tables for all parasites per species
Dichotomous table for leafminers
1a fleck mine; larva mines from a case => 2
1b corridor or blotch mine; larva mines from the inside => 3
1c galls, etc. => Tables for all parasites per species
2a case squat, hoary, greyish; mines only along the leaf margin: Coleophora pennella
2b case slender, almost bare, brown black; mines scattered over the leaf disk: Coleophora pulmonariella
3a larva with chitinised head; final mine at least partly full depth => 4
3b larva a maggot; mine never and nowhere full depth => 9
4a mines small, usually a large number in a leaf; larva without feet: Phytosciara macrotricha
4b mines usually single; larva with thoracic feet => 5
5a mines quite small; older larvae life free among spun leaves => 6
5b mines of normal size; larvae mine all their life => 7
6a pinacula colourless (but the bases of the setae themselves are black): Cnephasia incertana
6b pinacula black: C. stephensiana
7a blotch, preceded by a long narrow corridor => 8
7b blotch without a preceding corridor: Dialectica imperialella
8a pupa naked in the mine, upon pupation working itself halfway out: Dialectica scalariella
8b pupa in a cocoon, remains within the mine: Perittia echiella
9a long corridor; frass in discrete grains; puparium within the mine, in a, generally lower-surface, pupal chamber; the front spiracula of the puparium penetrate the epidermis: Chromatomyia horticola
9bmine different => 10
10a larva: cephalic skeleton with 3 backwards pointing arms; mine probably in all cases a primary blotch with a long initial corridor (may have been overrun by the blotch) => 11
10b cephalic skeleton with 3 backwards pointing arms; mine probably in all cases a blotch without a preceding corridor => 12 [this section of the key is based on Beiger (1975a)]
11a larva: mandible with 2 teeth, that alternate (seen from the side therefore virtually 4 teeth); mine communal, its first section already 5-10 mm wide: Agromyza ferruginosa
11b mandible with 2 teeth, not alternating; mine solitary, beginning with narrow preceding corridor (later the corridor may become overrun; moreover, mostly several blotches on a leaf fuse into one large and communal final mine): Agromyza abiens
11c the larvae of Agromyza myosotidis are unknown; species unknown from the Benelux (but that doesn’t mean much); probably they live solitary in not-coalescing blotches.
12a lavae communally => 13
12b larva solitary => 14
13a larva: front spiraculum with 17-19, rear spiraculum with 23-25 papillae: Phytomyza medicaginis
13b front spiraculum with 13-15, rear spiraculum with 19-21 papillae: Phytomyza nowakowskiana
14a larva: front spiraculum with 15-18, rear spiraculum with 18-19 papillae: Phytomyza myosotica [occurrence on Symphytum doubtfull]
14b front spiraculum with at most 10 papillae => 15
15a larva with big hump on the mesothorax: Phytomyza pulmonariae
15b no such hump => 16
16a mainly on Lithospermum: Phytomyza lithospermi [occurrence on Symphytum not quite certain]
16b on Symphytum ottomanum: Phytomyza rhodopaea [position in the key uncertain]
Not included in the key: Phytomyza obscura.