Phytomyza actaeae Hendel, 1922
on Actaea
Actaea spicata, GD Luxembourg, Hollenfels © Sébastien Carbonnelle
young mine and feeding punctures
mine
Large upper-surface, greyish-green, later brownish, blotch with conspicuous primary and secondary feeding lines. Often several larvae in a mine. Frass in irregularly dispersed granules. According to Hering (1957a), pupation takes place outside the mine, but Pakalniškis (2004a) states the opposite.
hostplants
Ranunculaceae, oligophagous
Actaea europaea, japonica, spicata.
Dreger & Myssura (2005a) additionally mention Clematis.
phenology
Mines from May to August (Hering, 1957a).
BENELUX
The picture above illustrates the first observation on the Benelux.
distribution within Europe
From Scandinavia to the Pyrenees, Alps, and Poland (Fauna Europaea, 2008); also Slovenia (Maček, 1999a).
larva
Mandibles with 2 alternating teeth; front spiraculum bifid, with ca. 12 papillae; rear spiraculum with 16-18 papillae (de Meijere, 1937a).
synonyms
Phytomyza nigrociliata Sasakawa, 1961.
notes
Also Beri (1971e) published a description of the larva. However, the hostplant he mentions, Aster asperulus, makes it clear that his identification is incorrect.
references
Ahr (1966a), Beiger (1955a, 1960a, 1965a), Beri (1971e), Buhr (1932a, 1941b, 1964a), Dreger & Myssura (2005a), Haase (1942a), Hering (1931f, 1955b, 1957a), Huber (1969a), Maček (1999a), de Meijere (1937a), Michna (1975a), Pakalniškis (2004a), Sasakawa (1961a), Skala & Zavřel (1945a), Sønderup (1949a), Spencer (1976a), Stammer (2016a), Starke (1942a), Starý (1930a), von Tschirnhaus (1999a).