Plant Parasites of Europe

leafminers, galls and fungi

Cynips divisa

Cynips divisa Hartig, 1840

red-pea gall

on Quercus, agamous generation

Cynips divisa gall

Quercus robur, Nieuwendam: gall, at underside of leaf

Cynips divisa gall

the wall is thck and hard …….

Cynips divisa gall

and consists of columnar cells

Cynips divisa gall

often the wall harbours one or two inquiline larvae ………

Cynips divisa gall

but they did not prevent this larva to reach the pupal stage

Cynips divisa pupa


Cynips divisa parasitoid

a parasitoid (with exuvium), attached to a larva


Galls from June till October, mature in September.

host plants

Fagaceae, monophagous

Quercus canariensis, frainetto, x hispanica, infectoria & subsp. veneris, lusitanica, macranthera, petraea, pubescens, pyrenaica, robur.

sexual generation

Cynips divisa gall

galls (from Houard, 1908a)


The sexual generation makes thin-walled, unilocular, lemon-shaped galls that with their tip are attached to a leaf margin, generally at the end on a vein. They are 2 x 3 mm in size, finely pubescent, and have about 10 length ridges. Galls also may be attached to the petiole of a catkin; these are larger, 4-6 mm long, blister-like and somewhat translucent.

host plants

Fagaceae, monophagous

Quercus frainetto, infectoria & subsp. veneris, petraea, pubescens, robur.

distribution within Europe

PESI (2024).


Dryophanta divisa; D. verrucosa (von Schlechtendal, 1870).


Of the practically unknown Cynips schlechtendali (Kieffer, 1901) it is stated by Houard that the gall is similar to the one of divisa, with the same finely granular surface, “but entirely smooth”.


Abras, Fassotte, Chandelier & Cavelier (2008a), Askew (1961b), Azmaz (2015a), Azmaz & Katilmiş (2017a), Bayrak (2019a), Béguinot (2002d,e, 2006a,c, 2007b), Bellmann (2012a), Blanes-Dalmau, Caballero-López & Pujade-Villar (2017a), Buhr (1965a), Cerasa (2015a), Chinery (2011a), Cogolludo (1921a), Coulianos & Holmåsen (1991a), Dauphin & Aniotsbehere (1997a), Demirel, Tataroğlu & Katilmiş (2023a), Eady & Quinlan (1963a), Ecott (2012a), Fusu (2017a), Groom (2011a), Hellrigl (2009a), Hellrigl & Bodur (2015a), Houard (1908a), Ilie & Marinescu (2011a), Karaca & Katılmış (2020a), Katılmış & Kıyak (2008a), Kemal & Koçak (2010a), Kollár (200a, 2011a), Koops (2013a), Kwast (2012a, 2014a), Lambinon, Carbonnelle & Claerebout (2015a), Lambinon, Schneider & Chevin (2003a), Lehmann & Flügel (2012a), Lehmann & Hannover (2016a), Melika (2006a, 2018a), Melika, Csóka & Pujade-Villar (2000a), Nieves-Aldrey (1984b, 2001a), Nieves-Aldrey, Gómez, Hernández Nieves & Lobo (2006a), Redfern & Shirley (2011a), Roskam (2009a), Shachar, Melika, Inbar & Dorchin (2018a), Tavares (1905a), Tomasi (2003a, 2014a), Williams (2010a).

Last modified 2.i.2024