Phyllonorycter harrisella (Linnaeus, 1761)
white oak midget

Quercus robur, Duin en Kruidberg

lighted from behind

Quercus robur, St Gerlach; opened mine
mine
Rather small, lower-surface tentiform mine, less than 14 mm long, with one strong fold in the lower epidermis. Pupa in a tough white cocoon that is attached to both the floor and roof of the mine. Before ecdysis the pupa half emerges from the underside of the mine.
host plants
Fagaceae, monophagous
Quercus Quercus cerris, dalechampii, faginea, frainetto, macranthera, petraea, pontica, pubescens, robur.
phenology
Larvae in July and September – October (Hering, 1957a).
BENELUX
BE recorded (Phegea, 2009).
NE recorded (Kuchlein & de Vos, 1999a; Microlepidoptera.nl, 2009).
LUX recorded (Fauna Europaea, 2009).
distribution within Europe
Europe, except the Balkan Peninsula and the Mediterranean Islands (Fauna Europaea, 2009).
pupa
synonyms
Lithocolletis harrisella; Glyphipteryx, Lithocolletis, Phyllonorycter cramerella (Fabricius, 1777).
parasitoids, predators
Achrysocharoides latreillii; Cirrospilus diallus; Euderus viridis.
notes
The biology is discussed by Miller (1973a).
According to Emmet ao (1985a) in the summer generation the frass is dispersed in the mine, and the cocoon is free of frass; in the autumn generation the frass is incrusted in the sides and rear of the cocoon, giving it a U-shaped outline when lighted from behind (see also Harper & Langmaid (1978a)). The picture above was made in mid-June and illustrates that in the Netherlands already in the summer generation almost all frass may be incorporated in the cocoon.
references
Bengtsson & Johansson (2011a), Buhr (1936a), Buszko (1992b), Corley, Marabuto & Pires (2007a), Corley, Rosete, Marabuto ao (2014a), Deutsch (2012a), Deutschmann (2008a), Emmet (1975b), Emmet, Watkinson & Wilson (1985a), van Frankenhuyzen Houtman & Kabos (1982a), ffennell (1975a), Gregor (1952a), Gregor & Patočka (2001a), Harper & Langmaid (1978a), Hartig (1939a), Hering (1934a, 1957a), Huemer (2012a), Huemer & Erlebach (2003a), Jaworski (2009a), Klimesch (1950c), Kuchlein & Donner (1993a), Kuchlein & de Vos (1999a), Lhomme (1934d), Maček (1999a), Matošević, Pernek, Dubravac & Barić (2009a), Miller (1973a), Nel & Varenne (2014a), Nowakowski (1954a), Patočka & Turčáni (2005a), De Prins, 1998a), Robbins (1991a), Schütze (1931a), Sefrová (2005a), Sønderup (1949a), Stolnicu (2007a), Szőcs (1977a, 1978a, 1981a), Tomov & Dimitrov (2007a), Tourlan (1980a), West (1985a), Yefremova & Kravchenko (2015a).