Heterodera humuli Filipjev, 1934
The roots often are abnormally short and strongly bushy branched. From mid-summer numerous globular bodies are attached, half a mm large, initially white, later yellowish brown to dark brown. They are cysts, the rear part of the females, filled with eggs. Inside the roots live numerous nematodes. Infected plants are strongly weakened.
Bongers (1988a), Buhr (1964b).