Plant Parasites of Europe

leafminers, galls and fungi

Pseudoperonospora cubensis

Pseudoperonospora cubensis (Berkeley & Curtis) Rostovzev, 1903

on Humulus, Cucurbitaceae

Pseudoperonospora cubensis on Humulus lupulus

Humulus lupulus, Belgium, prov. Antwerp, Balen, Griesbroek © Carina Van Steenwinkel

Pseudoperonospora cubensis on Humulus lupulus

infected leaf, upperside

Pseudoperonospora cubensis on Humulus lupulus


Pseudoperonospora cubensis on Humulus lupulus

underside of a leaf that has been infected longer ago

Pseudoperonospora cubensis: conidiophores


Pseudoperonospora cubensis: conidia



upperside of the leaves with angular, yellowish brown spots; at the underside a felty brownish purple fungal down, consisting of erect, distally strongly branched conidiophores. The conidia measure 10-20 x 14-32 µm. In particular young leaves may be strongly disfigured and diminutive in size.

host plants

Cannabaceae, Cucurbitaceae, polyphagous

Bryonia dioica; Cucumis melo, myriocarpus, sativus; Cucurbita pepo; Echinocystis lobata; Humulus lupulus; Sicyos angulatus.

To be expected on Bryonia dioica (Klenke & Scholler, 2015a).


Pseudoperonospora humuli (Miyabe & Takahashi) Wilson, 1914. Jage, Klenke, Kruse ao (2017a) discuss this as a valid species, with Humulus lupulus as a host plant.


Beenken & Senn-Irlet (2016a), Brandenburger (1985a: 74, 402), Buhr (1964b), Chater, Woods, Stringer, ao (2020a), Choi, Hong & Shin (2005a), Doppelbaur & Doppelbaur (1973a), Doppelbaur, Huber & Poelt (1965a), García-Blázquez, Constantinescu, Tellería & Martín (2007a), Göker, García-Blázquez, Voglmayr ao (2009a), Göker, Voglmayr, Riethmüller, Weiß & Oberwinkler (2003a), Gustavsson (1991a), Hafellner (1980a), Ilie & Marinescu (2011a), Jage, Klenke, Kruse ao (2016a, 2017a), Klenke & Scholler (2015a), Kozłowska, Mułenko & Heluta (2015a), Kruse (2019a), Kruse & Jage (2014a), Lebeda & Cohen (2011a), Müller (2015a), Müller & Kokeš (2008a), Mułenko, Piątek, Wołczańska ao (2010a), Negrean (1996b), Negrean & Anastasiu (2006a), Negrean, Constantinescu & Denchev (2004a), Porta-Puglia & Mifsud (2006a), Redfern & Shirley (2011a), Riethmüller, Voglmayr, Göker, Weiß & Oberwinkler (2002a), Ruszkiewicz-Michalska & Michalski (2005a), Savory, Granke, Quesada-Ocampo, ao (2011a), Scheuer & Bechter (2012a), Voglmayr (2003a).

Last modified 26.i.2023