Puccinia asperulina (Juel) Lagerheim, 1901
no host plant alternation. Aecia cupulate, numerous, often occupying the entire under surface of the leaf. Uredinia tiny, hypophyllous light brown.Telia similar but darker. Teliospores two-celled, obpyriform, wall thin and smooth; pedicel short, persistent. The fungus is systemic, and hibernates in the rhizome; infected shoots remain sterile and have an aberrant growth pattern.
Asperula ciliata, tinctoria.
Brandenburger (1985a), Buhr (1964b), Dauphin & Aniotsbehere (1997a), Gäumann (1959a), Klenke & Scholler (2015a), Poetl & Zwetko (1997a).