Puccinia cardui-pycnocephali Sydow & Sydow, 1902
on Carduus

Carduus bourgeanus, from González Fragoso (1924a): teliospores and two urediniospores
Carduus pycnocephalus: teliospore and an aberrant urediniospore
gall
no host plant alternation, only uredinia and telia. Uredinia dispersed, chestnut brown, pulverulent; urediniospores with 3-4 equatorial pores, each capped by a papilla. Telia also on the stem, dark brown, soon naked. Teliospores two-celled, elliptic, finely echinulate, c. 37 µm long. Pedicel hyaline, deciduous, up to 60 µm.
uredinia, telia
Asteraceae, narrowly monophagous
Carduus bourgeanus, pycnocephalus, tenuiflorus.
By Gäumann, and possibly on his authority also by Klenke & Scholler, “C. pycnocehalus” is interpreted as C. acanthoides; however, that is a different species.
synonyms
many authors, including Termorshuizen & Swertz, and also the Index Fungorum (2016), consider this species conspecific with P. calcitrapae.
references
Brandenburger (1985a), Gäumnn (1959a), González Fragoso (1924a), Klenke & Scholler (2015a), Kruse (2019a), Losa España (1944a), Termorshuizen & Swertz (2011a), Unamuno (1942a).