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on Dryopteris

Dryopteris spec., België, prov. Antwerpen, Balen, de Vennen © Carina Van Steenwinkel

uredinia

urediniospores

gall

uredinia hypophyllous, pustulate, covered with a brown epidermis, in which a pore develops; spores short club shaped, remotely spinulose, on a short, plump pedicel, c. 33 x 18 µm. From September on the uredinia produce a white mass of spores during damp weather. Telia at the underside of brown parts of the hibernated fronds. The spores are formed in the epidermis cells; they are irregular in shape, thin-walled, 1-40 celled.

host plants

Dryopteridaceae monophagous

Dryopteris x ambroseae, borreri, carthusiana, dilatata, expansa, filix-mas, intermedia subsp. azorica, pseudomas.

Termorshuizen & Swertz additionally mention Athyrium filix-femima (Woodsiaceae) and Polystichum aculeatum (Dryopteridaceae); on which this was based is not stated.

synonyms

Milesia kriegeriana.

references

Brandenburger (1985a: 6, 15), Bubner, Buchheit, Friedrich ao (2019a), Doppelbaur (1968a, 1973a), Eichhorn (1941a), Ellis & Ellis (1997a), Gäumann (1959a), Gjaerum & Dennis (1976a), Gjaerum & Sunding (1986a), González-Fragoso (1925a), Hafellner (1980a), Henderson (2004a), Hick & Preece (1990a), Jage, Klenke, Kruse ao (2016a, Klenke & Scholler (2015a), Kruse, Thiel, Choi ao (2016a), Poelt & Zwetko (1997a), Tănase & Negrean (2007a), Termorshuizen & Swertz (2011a), Tykhonenko & Hayova (2015a), Wilson & Henderson (1966a), Woods, Stringer, Evans & Chater (2015a).