Plant Parasites of Europe

leafminers, galls and fungi

Golovinomyces biocellatus

Golovinomyces biocellatus (Ehrenberg) Heluta, 1988

on Lamiaceae

Golovinomyces biocellatus: cleistothecia on Lycopus europaeus

Lycopus europaeus, Belgium, prov. Antwerp, Mol, Duivelskuil © Carina Van Steenwinkel

Golovinomyces biocellatus: appendages


Golovinomyces biocellatus: asci

two asci

Golovinomyces biocellatus: appressoria



mycelium amphigenous, mainly epiphyllous and along the main veins. Appressoria nipple-shaped, best developed in July and August. Conidia in short chains, without fibrosin bodies. Foot-cell of conidiophore 40-100 µm long, mostly distinctly curved. Cleistothecia with 5-15 asci, with 2 (up to 4) spores. Appendages numerous, mycelioid, septate, more or less brown, 0.5 – 2.5 x the cleistothecial diameter, attached to the lower half of the cleistothecium.

host plants

Lamiaceae, Nepetoideae, narrowly oligophagous:

Glechoma hederacea; Lycopus europaeus, exaltatus.


Kozłowska, Mułenko & Heluta (2015a), Kruse (2019a), Kruse, Thiel, Rätzel, ao (2020a), Leysen (2017a), Ruszkiewicz-Michalska & Michalski (2005a) , Scholler, Schmidt, Siahaan ao (2016a).

Last modified 7.iv.2023