Podosphaera macularis (Wallroth) Braun & Takamatsu, 2000
on Humulus, ? Cannabis

Humulus lupulus, Belgium, prov. Antwerp, Mol, 17.xi.2015 © Carina Van Steenwinkel

appendages numerous, brown …

… and septate

one single ascus …

… with up to eight spores
gall
mycelium amphigenous, white, patchy, persistent, may form a brownish crust when old. Conidia formed in chains, with fibrosin bodies. Cleistothecia often numerous, containing a single ascus with 4-8 spores. Appendages numerous, mostly sub-equatorial, 0.5-6 x the diameter; they are rather stiff and straight, unbranched, septate, brown.
host plants
Cannabaceae, monophagous
Humulus japonicus, lupulus.
According to Klenke & Scholler also, sterile, on Cannabis.
synonyms
Sphaerotheca humuli (de Candolle), Burrill, 1887; Sphaerotheca macularis (Wallroth) Magnus, 1899.
hyperparasites
Ampelomyces quisqualis; Mycodiplosis sphaerothecae.
references
Blumer (1946a, 1967a), Brandenburger (1985a: 75), Braun (1995a), Braun & Cook (2012a), Bresinsky (2016a), Butler & Jones (1955a), Czerniawska (2001a), Ellis & Ellis (1997a), Heluta, Hayova, Tykhonenko ao (2010a), Jage, Klenke & Kummer (2010a), Klenke & Scholler (2015a), Kozłowska, Mułenko & Heluta (2015a), Kruse (2019a), Leysen (2017a), Mayor (1967a), Redfern & Shirley (2011a), Roskam & Carbonnelle (2015a), Ruszkiewicz-Michalska & Michalski (2005a), Scholler & Schubert (1993a “Sphaerotheca spiraeae”), Tomasi (2014a), Tóth (1994a), Unamuno (1942a).