Plant Parasites of Europe

leafminers, galls and fungi

Podosphaera macularis

Podosphaera macularis (Wallroth) Braun & Takamatsu, 2000

on Humulus, ? Cannabis

Podosphaera macularis on Humulus lupulus

Humulus lupulus, Belgium, prov. Antwerp, Mol, 17.xi.2015 © Carina Van Steenwinkel

Podosphaera macularis: appendages

appendages numerous, brown …

Podosphaera macularis: septate appendage

… and septate

Podosphaera macularis: single ascus

one single ascus …

Podosphaera macularis: ascus

… with up to eight spores


mycelium amphigenous, white, patchy, persistent, may form a brownish crust when old. Conidia formed in chains, with fibrosin bodies. Cleistothecia often numerous, containing a single ascus with 4-8 spores. Appendages numerous, mostly sub-equatorial, 0.5-6 x the diameter; they are rather stiff and straight, unbranched, septate, brown.

host plants

Cannabaceae, monophagous

Humulus japonicus, lupulus.

According to Klenke & Scholler also, sterile, on Cannabis.


Sphaerotheca humuli (de Candolle), Burrill, 1887; Sphaerotheca macularis (Wallroth) Magnus, 1899.


Ampelomyces quisqualis; Mycodiplosis sphaerothecae.


Blumer (1946a, 1967a), Brandenburger (1985a: 75), Braun (1995a), Braun & Cook (2012a), Bresinsky (2016a), Butler & Jones (1955a), Czerniawska (2001a), Ellis & Ellis (1997a), Heluta, Hayova, Tykhonenko ao (2010a), Jage, Klenke & Kummer (2010a), Klenke & Scholler (2015a), Kozłowska, Mułenko & Heluta (2015a), Kruse (2019a), Leysen (2017a), Mayor (1967a), Redfern & Shirley (2011a), Roskam & Carbonnelle (2015a), Ruszkiewicz-Michalska & Michalski (2005a), Scholler & Schubert (1993a “Sphaerotheca spiraeae”), Tomasi (2014a), Tóth (1994a), Unamuno (1942a).

Last modified 3.xii.2021