Endophyllum sempervivi (Albertini & Schweinitz) de Bary, 1863
on Sempervivum

Sempervivum tectorum, Belgium, prov. Liège, Ferrières, 20.iv.2012 © Jean-Yves Baugnée

detail with telia (here as small pits)

Sempervivum spec., Italy, Aosta, vii.2013 © Ben van As
Aeonium spec., Canary IsLlands, Tenerife, RN Teno, 9.ii.2020 © Sébastien Carbonelle
detail
gall
Endophyllum species have but two spore stages: spermogonia and aecia. The spermogonia are largely sunken into the leaf tissue, only protruding as small, brown cones. Infested leaves often are strongly elongated.
host plants
Crassulaceae, oligophagous
Echeveria; Jovibarba globifera & subsp. arenaria + hirta; Sempervivum arachnoideum, calcareum, montanum, tectorum, wulfenii.
As demonstrated by the pictures above, this, or a closely related, species also occurs on Aeonium.
notes
Can destroy the plant. Only the leaves that have been formed in early summer are affected. Diseased leaves are highly attractive to aphids.
references
Bahcecioglu & Kabaktepe (2012a), Bellmann (2012a), Blumer (1946a), Buhr (1965a), Coulianos & Holmåsen (1991a), Dauphin & Aniotsbehere (1997a), Gäumann (1959a), González-Fragoso (1925a), Henderson (2000a, 2004a), Jage, Kruse, Kummer ao (2013a), Klenke & Scholler (2015a), Kruse (2019a), Ljaljević Grbić, Lakušić, Stupar & Vukojević (2016a), Llorens i Villagrasa (1984a), Maier, Wingfield, Mennicken & Wingfield (2007a), Poelt & Zwetko (1997a), Preece & Hick (1994a), Redfern & Shirley (2011a), Roskam (2009a), Ruszkiewicz-Michalska (2006a), Scheuer & Bechter (2012a), Termorshuizen & Swertz (2011a), Unamuno (1941a), Vanderweyen & Fraiture (2008a), Zwetko (2000a).