Puccinia betonicae (Albertini & Schweinitz) de Candolle, 1815
on Stachys
Stachys officinalis, Belgium, prov. Namur, Parc National de Furfooz © Stéphane Claerebout: a normal leaf and an infected one
telia

Stachys officinalis, Belgium, prov. Namur, Honnai; leg. J-M Couvreur, det. Arthur Vanderweyen © Jean-Yves Baugnée
Stachys officinalis, from González Fragoso (1924a): teliospores
gall
No host plant alternation, only telia. The mycelium is systemic; infected leaves are pale and have a long petiole; the plants get an abnormal aspect. Telia mostly hypophyllous, brown, pulverulent; spores two-celled, smooth, on a short deciduous pedicel.
host plants
Lamiaceae, narrowly monophagous
Stachys annua, citrina, cretica, iberica, macrantha, officinalis & subsp. serotina, pradica.
references
Bahçecioğlu & Kabaktepe (2012a), Bahçecioğlu, Kabaktepe & Yildiz (2006a), Brandenburger (1985a: 543), Buhr (1965a), Dauphin & Aniotsbehere (1997a), Ellis & Ellis (1997a), Gäumann (1959a), González Fragoso (1924a), Henderson (2000a, 2004a), Jage, Kruse, Kummer ao (2013a), Klenke & Scholler (2015a), Llorens i Villagrasa (1984a), Poelt & Zwetko (1997a), Redfern & Shirley (2011a), Termorshuizen & Swertz (2011a), Tomasi (2012a, 2014a), Vanderweyen & Fraiture (2011a), Wilson & Henderson (1966a), Woods, Stringer, Evans & Chater (2015a).